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Modern humans have neither an air sac nor an extension on the hyoid bone.
But —a hominin species that roamed Africa approximately 3.9 million to 2.9 million years ago—had a hyoid bulla, the fossil record shows, meaning it’s highly likely it had an air sac, too.
Non-human primates have an organ called an air sac, a large cavity that connects to the vocal tract.
The air sac links onto an extension on the hyoid bone known as the hyoid bulla.
Using plastic tubing, de Boer built models of the human vocal tract both without an air sac, like modern humans, and with one, like The air sacs acted like bass drums, resonating at low frequencies, and causing vowel sounds to merge; [an Australopithecus] would have had a greatly reduced vocabulary.